Amphibia-Reptilia 5 (1984): 69-78

Summer Activity Patterns of Testudo hermanni GMELIN in Greece and France

Adrian Hailey1, Elizabeth A. Pulford2, and David Stubbs3

1 Department of Zoology, University of Nottingham, NG7 2RD, U.K.
2 Department of Zoology, University College of North Wales, Bangor, Gwynedd LL57 2UW, U.K.
3 School of Continuing Education, Rutherford College, University of Kent, Canterbury CT2 7NX, U.K.

Abstract. Daily activity patterns of Testudo hermanni in summer are described for two sites, Alyki, a coastal heathland in Greece, and the Massif des Maures, a woodland in France. Activity was bimodal in the former and unimodal in the latter. Habitat type was thought to be more important than latitude and climate in accounting for this pattern. At Alyki there was a sex ratio of encounters of 2.7 males : 1 female in summer, but no differences between the sexes in daily activity pattern. In the Massif des Maures, the ratio varied through the season, from 0.5 male : 1 female in June, to 2.6 males : 1 female in September, suggesting seasonal differences in activity of the sexes. Population density at Alyki was 5 times greater than in the Massif des Maures, yet sighting frequency was similar. This was accounted for by the increased ease of finding active animals in the woodland from the noise they made in the leaf litter.